Professor Marysia Placzek

School of Biosciences

Professor of Developmental Neurobiology

m.placzek@sheffield.ac.uk
Firth Court

Full contact details

Professor Marysia Placzek
School of Biosciences
Firth Court
Western Bank
Sheffield
S10 2TN
Profile

Marysia is a Wellcome Trust Investigator and Professor of Developmental Neurobiology in the School of Biosciences. She has inspired staff and students alike with her pioneering leadership.

A driving force behind one of our University’s key research facilities, Marysia directed the MRC Centre for Developmental and Biomedical Genetics for many years and oversaw its evolution into the Bateson Centre.

Her initiative and outstanding commitment to her colleagues and students contributed to the ongoing success of research in the Department and the strong focus on developmental biology, and its translation, at the University.

Marysia’s research focuses on the development of the vertebrate hypothalamus and she has earned international acclaim for her work in this field. In 2023 she was awarded the British Society for Developmental Biology Waddington Medal, in recognition of her outstanding research performance as well as services to the subject community.

In 2012 she was awarded an MRC Suffrage Science Heirloom in recognition of her achievements as a leading female scientist.

Like many women, she balances the demands of world-leading research with family commitments, and has been celebrated by The Royal Society’s ‘Parent Carer Scientist’ project.

Colleagues agree that “Marysia is not just a role model for female scientists, but for all those with an aspiration to become a leader in their field.”

Research interests
Chick hyp

How the brain develops remains one of the greatest mysteries in science. Research in the lab focus on the development of a fascinating part of the brain - the hypothalamus – the master-controller of physical, behavioural and emotional homeostasis.

Lay summary

In the developing embryos, the hypothalamus is built with precision. This ordered assembly underlies its key role in life. Our research focuses on characterising the stem and progenitor cells that build the hypothalamus and characterising the molecular networks that direct hypothalamic morphogenesis, growth and differentiation. Our work will contribute to understanding the importance of the hypothalamus to robust long-term health and will shed light on diseases and disruptions of homeostasis.

Research summary

The functions of the hypothalamus in mediating homeostasis, and ensuring that brain and body function optimally, are well-known. By contrast, little is understood of how the hypothalamus develops. This knowledge is important, because early indications suggest that deregulation of developmental programmes may underlie complex human pathological conditions, including stress and eating disorders. Our goal is therefore to understand how the hypothalamus develops in the embryo and how the proper embryonic assembly of the hypothalamus holds the key to robust adult function.

Chick

We focus in particular on five key areas:

  • The role of adjacent tissues in inducing multipotent embryonic hypothalamic progenitor cells
  • The characterisation of multipotent embryonic hypothalamic progenitor cells, and identification of cues that maintain progenitor cells, or promote their differentiation
  • The self-organisation and morphogenesis of hypothalamic progenitor cells through integrated growth and differentiation
  • The cellular and molecular events that underlie the integrated assembly of the hypothalamo-pituitary neuraxis
  • The characterization of stem/progenitor-like tanycytes in the adult hypothalamus

We use a range of animal model systems (chick, mouse, zebrafish) and combine in vivo and ex vivo approaches with state-of-the-art transcriptomic, imaging, and gain-and loss-of-function approaches.

Tanycytes
Publications

Show: Featured publications All publications

Journal articles

  • Groves I, Holmshaw J, Furley D, Manning E, Chinnaiya K, Towers M, Evans BD, Placzek M & Fletcher AG (2023) . Development, 150(22).
  • Chinnaiya K, Burbridge S, Jones A, Kim DW, Place E, Manning E, Groves I, Sun C, Towers M, Blackshaw S & Placzek M (2023) . eLife, 12.
  • Kim DW, Place E, Chinnaiya K, Manning E, Sun C, Dai W, Groves I, Ohyama K, Burbridge S, Placzek M & Blackshaw S (2022) . Cell Reports, 38(3).
  • Brown SJ, Boussaad I, Jarazo J, Fitzgerald JC, Antony P, Keatinge M, Blechman J, Schwamborn JC, Krüger R, Placzek M & Bandmann O (2021) . Scientific Reports, 11(1).
  • Tait CM, Chinnaiya K, Manning E, Murtaza M, Ashton J-P, Furley N, Hill CJ, Alves CH, Wijnholds J, Erdmann KS , Furley A et al (2020) . PLOS Biology, 18(3).
  • Frith TJR, Granata I, Wind M, Stout E, Thompson O, Neumann K, Stavish D, Heath PR, Ortmann D, Hackland JOS , Anastassiadis K et al (2018) . eLife, 7.
  • Placzek M & Briscoe J (2018) . International Journal of Developmental Biology, 62(1-3), 221-230.
  • Fu T, Towers M & Placzek M (2017) . Development, 144(18), 3278-3288.
  • Burbridge S, Stewart I & Placzek M (2016) . Comprehensive Physiology, 6(2), 623-643.
  • Robins SC, Stewart I, McNay DE, Taylor V, Giachino C, Goetz M, Ninkovic J, Briancon N, Maratos-Flier E, Flier JS , Kokoeva MV et al (2013) . Nat Commun, 4, 2049.

Book chapters

  • Placzek M, Fu T & Towers M (2020) , Masterclass in Neuroendocrinology (pp. 3-30). Springer International Publishing

Preprints

  • Kim DW, Place E, Chinnaiya K, Manning E, Sun C, Dai W, Ohyama K, Burbridge S, Placzek M & Blackshaw S (2021) , Cold Spring Harbor Laboratory.
  • Tait C, Chinnaiya K, Murtaza M, Ashton J-P, Furley N, Hill CJ, Henrique Alves C, Wijnholds J, Erdmann KS, Das RM , Rashbass P et al (2019) , Cold Spring Harbor Laboratory.

All publications

Journal articles

  • Chinnaiya K, Groves I, Manning E, Place E, Furley C, Strawbridge SE, Fletcher AG & Placzek M (2025) . Developmental Biology, 525, 259-269.
  • Placzek M, Chinnaiya K, Kim DW & Blackshaw S (2025) . Nature Reviews Endocrinology, 21(2), 118-130.
  • Beal S, Stewart I, Hatton P, Placzek M & Ortega I (2024) . Engineered Regeneration, 5(1), 11-20.
  • Chinnaiya K & Placzek M (2023) . BIO-PROTOCOL, 13(23).
  • (2023) . Development, 150(23).
  • Groves I, Holmshaw J, Furley D, Manning E, Chinnaiya K, Towers M, Evans BD, Placzek M & Fletcher AG (2023) . Development, 150(22).
  • Eachus H, Oberski L, Paveley J, Bacila I, Ashton J-P, Esposito U, Seifuddin F, Pirooznia M, Elhaik E, Placzek M , Krone N et al (2023) . Disease Models & Mechanisms, 16(9).
  • Sedas Perez S, McQueen C, Stainton H, Pickering J, Chinnaiya K, Saiz-Lopez P, Placzek M, Ros MA & Towers M (2023) . Nature Communications, 14(1).
  • Chinnaiya K, Burbridge S, Jones A, Kim DW, Place E, Manning E, Groves I, Sun C, Towers M, Blackshaw S & Placzek M (2023) . eLife, 12.
  • Eachus H, Ryu S, Placzek M & Wood J (2022) . Current Opinion in Endocrine and Metabolic Research, 26.
  • Moore A, Chinnaiya K, Kim DW, Brown S, Stewart I, Robins S, Dowsett GKC, Muir C, Travaglio M, Lewis JE , Ebling F et al (2022) . Frontiers in Neuroscience, 16.
  • Kim DW, Place E, Chinnaiya K, Manning E, Sun C, Dai W, Groves I, Ohyama K, Burbridge S, Placzek M & Blackshaw S (2022) . Cell Reports, 38(3).
  • Wind M, Gogolou A, Manipur I, Granata I, Butler L, Andrews PW, Barbaric I, Ning K, Guarracino MR, Placzek M & Tsakiridis A (2021) . Development, 148(6).
  • Brown SJ, Boussaad I, Jarazo J, Fitzgerald JC, Antony P, Keatinge M, Blechman J, Schwamborn JC, Krüger R, Placzek M & Bandmann O (2021) . Scientific Reports, 11(1).
  • De Souza FSJ & Placzek M (2021) . The International Journal of Developmental Biology, 65(4-5-6), 195-205.
  • Jones A, Barker‐Haliski M, Ilie AS, Herd MB, Baxendale S, Holdsworth CJ, Ashton J, Placzek M, Jayasekera BAP, Cowie CJA , Lambert JJ et al (2020) . Epilepsia, 61(10), 2106-2118.
  • Groves I, Placzek M & Fletcher AG (2020) . Philosophical Transactions of the Royal Society B: Biological Sciences, 375(1809).
  • Tait CM, Chinnaiya K, Manning E, Murtaza M, Ashton J-P, Furley N, Hill CJ, Alves CH, Wijnholds J, Erdmann KS , Furley A et al (2020) . PLOS Biology, 18(3).
  • Fu T, Pearson C, Towers M & Placzek M (2019) . Journal of Neuroendocrinology, 31(5).
  • Eachus H, Bright C, Cunliffe VT, Placzek M, Wood JD & Watt PJ (2018) . ESPE Yearbook of Paediatric Endocrinology, 15.
  • Frith TJR, Granata I, Wind M, Stout E, Thompson O, Neumann K, Stavish D, Heath PR, Ortmann D, Hackland JOS , Anastassiadis K et al (2018) . eLife, 7.
  • Placzek M & Briscoe J (2018) . International Journal of Developmental Biology, 62(1-3), 221-230.
  • Fu T, Towers M & Placzek M (2017) . Development, 144(18), 3278-3288.
  • Eachus H, Bright C, Cunliffe VT, Placzek M, Wood JD & Watt PJ (2017) . Human Molecular Genetics, 26(11), 1992-2005.
  • Muthu V, Eachus H, Ellis P, Brown S & Placzek M (2016) . Development, 143(14), 2651-2663.
  • Burbridge S, Stewart I & Placzek M (2016) . Comprehensive Physiology, 6(2), 623-643.
  • Burbridge S, Stewart I & Placzek M (2016) . Comprehensive Physiology, 6(2), 623-643.
  • Ellis PS, Burbridge S, Soubes S, Ohyama K, Ben-Haim N, Chen C, Dale K, Shen MM, Constam D & Placzek M (2015) . Development, 142(22), 3821-3832.
  • Liua F, Placzekb M & Xu H (2014) . Neuroscience Letters, 562, 107.
  • Liu F & Placzek M (2013) . Neuroscience Letters.
  • Christou YA, Ohyama K, Placzek M, Monk PN & Shaw PJ (2013) . BMC Neurosci, 14, 126.
  • Liu F, Placzek M & Xu H (2013) . Neurosci Lett, 553, 104-109.
  • Robins SC, Stewart I, McNay DE, Taylor V, Giachino C, Goetz M, Ninkovic J, Briancon N, Maratos-Flier E, Flier JS , Kokoeva MV et al (2013) . Nat Commun, 4, 2049.
  • Liu F, Pogoda H-M, Pearson CA, Ohyama K, Löhr H, Hammerschmidt M & Placzek M (2013) . Development, 140(5), 1111-1122.
  • Pearson CA & Placzek M (2013) . Curr Top Dev Biol, 106, 49-88.
  • Placzek M & Anton ES (2013) . CELL STEM CELL, 12(1), 10-11.
  • Pearson CA, Ohyama K, Manning L, Aghamohammadzadeh S, Sang H & Placzek M (2011) . Development, 138(12), 2613-2624.
  • Van Hateren NJ, Das RM, Hautbergue GM, Borycki A-G, Placzek M & Wilson SA (2011) . Development, 138(10), 1893-1902.
  • Blackshaw S, Scholpp S, Placzek M, Ingraham H, Simerly R & Shimogori T (2010) . J Neurosci, 30(45), 14925-14930.
  • Placzek M (2008) . Methods Mol Biol, 461, 325-335.
  • Ohyama K, Das R & Placzek M (2008) . Development, 135(20), 3325-3331.
  • Manning L, Ohyama K, Saeger B, Hatano O, Wilson SA, Logan M & Placzek M (2006) . Dev Cell, 11(6), 873-885.
  • Ingham PW & Placzek M (2006) . NAT REV GENET, 7(11), 841-850.
  • Das RM, Van Hateren NJ, Howell GR, Farrell ER, Bangs FK, Porteous VC, Manning EM, McGrew MJ, Ohyama K, Sacco MA , Halley PA et al (2006) . Dev Biol, 294(2), 554-563.
  • Ohyama K, Ellis P, Kimura S & Placzek M (2005) . Development, 132(23), 5185-5197.
  • Chu J, Ding J, Jeays-Ward K, Price SM, Placzek M & Shen MM (2005) . Development, 132(24), 5539-5551.
  • Placzek M & Briscoe J (2005) . Nat Rev Neurosci, 6(3), 230-240.
  • Jeays-Ward K & Placzek M (2005) Molecular control of area a cell specification to anterior floor plate fate. MECHANISMS OF DEVELOPMENT, 122, S164-S165.
  • Pearson CA, Manning L, Ohyama K & Placzek M (2005) Neurogenesis in the chick hypothalamus. MECHANISMS OF DEVELOPMENT, 122, S177-S177.
  • Pevny L & Placzek M (2005) . Curr Opin Neurobiol, 15(1), 7-13.
  • Patten I, Kulesa P, Shen MM, Fraser S & Placzek M (2003) . Development, 130(20), 4809-4821.
  • Patten I & Placzek M (2002) . Curr Biol, 12(1), 47-52.
  • Patten I & Placzek M (2001) . Curr Biol, 11(15), R616-R619.
  • Ellis S, Patten I, Vesque C, Shen M, Doddì J & Placzek M (2000) . Biochemical Society Transactions, 28(5), A139-A139.
  • Vesque C, Ellis S, Lee A, Szabo M, Thomas P, Beddington R & Placzek M (2000) . Development, 127(13), 2795-2809.
  • Placzek M & Ingham PW (2000) Developmental genetics in Sheffield: a meeting point for Hedgehog researchers.. Int J Dev Biol, 44(1), 65-72.
  • Placzek M, Dodd J & Jessell TM (2000) . Curr Opin Neurobiol, 10(1), 15-22.
  • Soubes SC & Placzek M (2000) Rostro-ventral patterning of the neural tube ventral midline. EUROPEAN JOURNAL OF NEUROSCIENCE, 12, 12-12.
  • Patten I & Placzek M (2000) . Cell Mol Life Sci, 57(12), 1695-1708.
  • Ingham PW & Placzek M (2000) Introduction: preface to the Hedgehog family of proteins review volume. CELL MOL LIFE SCI, 57(12), 1671-1671.
  • Placzek M & Skaer H (1999) . Curr Biol, 9(14), R506-R510.
  • Dale K, Sattar N, Heemskerk J, Clarke JD, Placzek M & Dodd J (1999) . Development, 126(2), 397-408.
  • Placzek M & Dale K (1999) . Methods Mol Biol, 97, 293-304.
  • Tessier-Lavigne M & Placzek M (1998) . Principles of Medical Biology, 11(C), 153-165.
  • Dodd J, Jessell TM & Placzek M (1998) . Science, 282(5394), 1654-1657.
  • Pevny LH, Sockanathan S, Placzek M & Lovell-Badge R (1998) . Development, 125(10), 1967-1978.
  • Dale JK, Vesque C, Lints TJ, Sampath TK, Furley A, Dodd J & Placzek M (1997) . Cell, 90(2), 257-269.
  • Placzek M & Furley A (1996) . Curr Biol, 6(5), 526-529.
  • Ericson J, Muhr J, Placzek M, Lints T, Jessell TM & Edlund T (1995) . Cell, 81(5), 747-756.
  • ERICSON J, MUHR J, PLACZEK M, LINTS T, JESSELL TM & EDLUND T (1995) SONIC HEDGEHOG INDUCES THE DIFFERENTIATION OF VENTRAL FOREBRAIN NEURONS - A COMMON SIGNAL FOR VENTRAL PATTERNING WITHIN THE NEURAL-TUBE (VOL 81, PG 747, 1995). CELL, 82(1), U11-U11.
  • Ruiz i Altaba A, Placzek M, Baldassare M, Dodd J & Jessell TM (1995) . Dev Biol, 170(2), 299-313.
  • Placzek M (1995) . Curr Opin Genet Dev, 5(4), 499-506.
  • Roelink H, Augsburger A, Heemskerk J, Korzh V, Norlin S, Ruiz i Altaba A, Tanabe Y, Placzek M, Edlund T & Jessell TM (1994) . Cell, 76(4), 761-775.
  • Placzek M, Jessell TM & Dodd J (1993) . Development, 117(1), 205-218.
  • Wilson SW, Placzek M & Furley AJ (1993) . Trends Neurosci, 16(8), 316-323.
  • Placzek M, Yamada T, Tessier-Lavigne M, Jessell T & Dodd J (1991) Control of dorsoventral pattern in vertebrate neural development: induction and polarizing properties of the floor plate.. Dev Suppl, Suppl 2, 105-122.
  • Yamada T, Placzek M, Tanaka H, Dodd J & Jessell TM (1991) . Cell, 64(3), 635-647.
  • Tessier-Lavigne M & Placzek M (1991) . Trends Neurosci, 14(7), 303-310.
  • Placzek M, Tessier-Lavigne M, Yamada T, Jessell T & Dodd J (1990) . Science, 250(4983), 985-988.
  • Placzek M, Tessier-Lavigne M, Jessell T & Dodd J (1990) . Development, 110(1), 19-30.
  • Placzek M, Tessier-Lavigne M, Yamada T, Dodd J & Jessell TM (1990) . Cold Spring Harb Symp Quant Biol, 55, 279-289.
  • Jessell TM, Bovolenta P, Placzek M, Tessier-Lavigne M & Dodd J (1989) . Ciba Found Symp, 144, 255-276.
  • Peters G, Brookes S, Smith R, Placzek M & Dickson C (1989) . Proc Natl Acad Sci U S A, 86(15), 5678-5682.
  • Peters G, Brookes S, Placzek M, Schuermann M, Michalides R & Dickson C (1989) . J Virol, 63(3), 1448-1450.
  • Tessier-Lavigne M, Placzek M, Lumsden AG, Dodd J & Jessell TM (1988) . Nature, 336(6201), 775-778.
  • Brookes S, Placzek M, Moore R, Dixon M, Dickson C & Peters G (1986) . Nucleic Acids Res, 14(21), 8231-8245.
  • Peters G, Placzek M, Brookes S, Kozak C, Smith R & Dickson C (1986) . J Virol, 59(3), 535-544.

Book chapters

  • Manning E & Placzek M (2024) , Current Topics in Developmental Biology (pp. 83-123). Elsevier
  • Placzek M, Fu T & Towers M (2020) , Masterclass in Neuroendocrinology (pp. 3-30). Springer International Publishing
  • Albertine KA, Alvarez IS, Bally-Cuif L, Barald KF, Bersu E, Brothman A, Byrne JLB, Capel B, Carey JC, Chen Y , Edlund H et al (2009) , Larsen's Human Embryology (pp. v-v). Elsevier
  • Placzek M (2006) , Imaging the Central Nervous System of the Fetus and Neonate (pp. 3-12). CRC Press
  • Jessell TM, Bovolenta P, Karagogeos D, Placzek M, Tessier-Lavigne M & Dodd J (1990) , Brain Repair (pp. 199-211). Macmillan Education UK

Conference proceedings

  • Bandmann S, Keatinge M, Placzek M & Bandmann O (2018) . Movement Disorders, Vol. 33(S2) (pp S605-S605). Hong Kong, China
  • Stokes S, Mani N, Bahl A, O'Connor-Read L & Placzek M (2013) Adult stem cells in the human and mouse spinal cord: What are their characteristics and can they migrate and replicate upon exposure to growth factors. BRITISH JOURNAL OF SURGERY, Vol. 100 (pp 38-39)
  • Hoggar EC, Placzek M & Chico TJ (2012) . HEART, Vol. 98 (pp A4-A4)
  • Hoggar EC, Placzek M & Chico TJ (2011) . HEART, Vol. 97(20) (pp 15-15)
  • Hoggar E, Pugh S, Wilson S, Placzek M, Crossman D & Chico T (2009) . HEART, Vol. 95(22)
  • Placzek M (2009) . MECHANISMS OF DEVELOPMENT, Vol. 126 (pp S8-S8)
  • Ohyama K, Wilson S, Briscoe J & Placzek M (2009) . MECHANISMS OF DEVELOPMENT, Vol. 126 (pp S210-S210)
  • Muthu V & Placzek M (2009) . MECHANISMS OF DEVELOPMENT, Vol. 126 (pp S220-S220)
  • Parkin C, Baxendale S, Smith S & Placzek M (2009) . MECHANISMS OF DEVELOPMENT, Vol. 126 (pp S142-S142)
  • Pearson CA, Ohyama K & Placzek M (2009) . DEVELOPMENTAL BIOLOGY, Vol. 331(2) (pp 467-468)
  • Hoggar EC, Pugh SD, Wilson S, Placzek M, Crossman DC & Chico TJA (2009) A NOVEL CHICK EMBRYO MODEL REVEALS ENDOTHELIN RECEPTOR B IS ESSENTIAL FOR COLLATERAL VESSEL DEVELOPMENT. HEART, Vol. 95 (pp A65-A65)
  • Manning L, Ohyama K, Soubes S & Placzek M (2003) A role for BMPs in the formation of the ventral hypothalamus.. DEVELOPMENTAL BIOLOGY, Vol. 259(2) (pp 492-492)

Preprints

  • Manning E, Chinnaiya K, Furley C, Kim DW, Blackshaw S, Placzek M & Place E (2025) , Cold Spring Harbor Laboratory.
  • Chinnaiya K, Burbridge S, Jones A, Kim DW, Place E, Manning E, Groves I, Sun C, Towers M, Blackshaw S & Placzek M (2022) , Cold Spring Harbor Laboratory.
  • Groves I, Holmshaw J, Furley D, Towers M, Evans BD, Placzek M & Fletcher AG (2022) , Cold Spring Harbor Laboratory.
  • Moore A, Chinnaiya K, Kim DW, Brown S, Stewart I, Robins S, Dowsett G, Muir C, Travaglio M, Lewis JE , Ebling F et al (2021) , Cold Spring Harbor Laboratory.
  • Kim DW, Place E, Chinnaiya K, Manning E, Sun C, Dai W, Ohyama K, Burbridge S, Placzek M & Blackshaw S (2021) , Cold Spring Harbor Laboratory.
  • Wind M, Gogolou A, Manipur I, Granata I, Butler L, Andrews PW, Barbaric I, Ning K, Guarracino MR, Placzek M & Tsakiridis A (2020) , Cold Spring Harbor Laboratory.
  • Tait C, Chinnaiya K, Murtaza M, Ashton J-P, Furley N, Hill CJ, Henrique Alves C, Wijnholds J, Erdmann KS, Das RM , Rashbass P et al (2019) , Cold Spring Harbor Laboratory.
  • Eachus H, Subramanya D, Jackson HE, Wang G, Berntsen K, Ashton J-P, Esposito U, Seifuddin F, Pirooznia M, Elhaik E , Krone N et al (2019) , Cold Spring Harbor Laboratory.
  • Kim DW, Place E, Chinnaiya K, Manning E, Sun C, Dai W, Ohyama K, Burbridge S, Placzek M & Blackshaw S () .
Research group

Current lab members:

  • Dr Sarah Burbridge (Research Assistant)
  • Dr Kavitha Chinnaiya (Research Associate)
  • Dr Elizabeth Manning (Research Associate)
  • Dr Elsie Place (Research Associate)
  • Mr Ian Groves (PhD student; joint with Dr Alex Fletcher)
  • Ms Bethany James (PhD student; joint with Professor Andrew Furley and Dr Ivana Barbaric)
Teaching activities

I lecture and teach on several undergraduate and MSc courses, including second, third year and masters courses in developmental biology, developmental neurobiology and stem/ regenerative biology. I am module co-ordinator for Developmental Neurobiology (second year module).

My group annually takes several undergraduate and masters students into the lab for their final dissertation/research projects and most years we have summer students working with us.

I am an invited editor of ‘Principles of Development Biology’ (7th ed).

Professional activities and memberships
  • Chair: Wellcome Trust Expert Review Group: Cell and Developmental Biology
  • Scientific Advisory Board member: GW4 Biomed MRC DTP

Presentations

I am invited to present research talks at international meetings in numerous fields, including Developmental biology, Neuroscience, Neuroendocrinology, Endocrinology

Examination bodies

  • External Examiner for Development, Regeneration and Stem Cells, University of Edinburgh (2016-2019)
  • External Examiner for Part 2 Zoology, University of Cambridge (2013-2015)

Professional societies

  • British Society for Developmental Biology
  • Society for Endocrinology
  • Society for Neuroscience
  • Society for Anatomy

Peer review activity

I am an ad hoc reviewer for many journals (including Cell, Nature Neuroscience, Current Biology, Development, Developmental Cell, PLoS Biology) and for Grant awarding bodies (including Wellcome Trust, BBSRC, MRC, ERC, Hong Kong Research Council, Human Science Frontiers Program; NIHR). I have been involved in many promotion reviews for scientists in my field of research in the UK, USA and elsewhere, act as a Content Expert (eg Larsen’s Human Embryology 4th Ed (Ed: G. Schoenwolf)) and editor (Wolpert’s Principlels of Developmental Biology 7th Ed).

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